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Family Miridae - Plant Bugs

Garden Fleahopper - Microtechnites bractatus - female Plant Bug - Miridae? - Phytocoris nigricollis Mirid I - Plagiognathus Elongate bug - mirid?  Stenodemini? - Leptopterna Mirid - Neolygus Tiny bug on Tall Thistle Seed-Plumes - Lygus Colorful plant bug - Phytocoris Male, Taedia marmorata? - male
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Heteroptera (True Bugs)
Infraorder Cimicomorpha
Superfamily Miroidea
Family Miridae (Plant Bugs)
Other Common Names
Plant bugs, Leaf bugs
Synonyms and other taxonomic changes
Miridae Hahn 1833 (sometimes listed as 1831)
close to 2,000 species in 223 genera in our area(1); the largest heteropteran family, with >10,000 described species in ~1,300 genera worldwide(2)(3)
adults 1.5-15 mm (more in some neotropical Restheniini)(3)
The unique characteristic of mirids is special setae ('trichobothria') on middle and hind femora
Other useful features:
lack of small, simple eyes called ocelli (NB: ocelli may be absent in other bugs, e.g. Cnemodus or Largidae, yet present in a few mirids (Isometopinae)
presence of a cuneus (a hind section of the thickened anterior part of the front wing, set off from it by a crease) - also present in Anthocoridae, Cimicidae, and Microphysidae
antennae mostly long and thin (usually shorter/thicker in Lygaeoidea and Coreoidea)
legs slender, delicate
most are plant feeders, but many are predatory or zoophytophagous
Overview of our fauna(4):
Subfamily Bryocorinae
Tribe Bryocorini

Tribe Dicyphini
Genera Campyloneura, Cyrtopeltis, Dicyphus, Engytatus, Macrolophus, Miomonalonion, Nesidocoris, Tupiocoris, Usingerella

Genera Agaveocoris, Cyrtocapsus, Halticotoma, Hesperolabops, Pycnoderes, Pycnoderella, Sixeonotopsis, Sixeonotus, Sysinas

Subfamily Cylapinae
Tribe Cylapini
Genera Cylapus

Tribe Fulviini

Subfamily Deraeocorinae
Tribe Clivinemini
Genera Ambracius, Bothynotus, Clivinema, Largidea

Tribe Deraeocorini
Genera Conocephalocoris, Deraeocapsus, Deraeocoris, Diplozona, Eurychilopterella, Hesperophylum, Klopicoris, Strobilocapsus(5)

Tribe Hyaliodini
Genera Hyaliodes, Paracarniella, Stethoconus

Tribe Surinamelini
Genera Eustictus

Subfamily Isometopinae
Tribe Diphlebini
Genera Diphleps

Tribe Myiommini
Genera Corticoris, Lidopus, Myiomma, Wetmorea

Subfamily Mirinae
Tribe Herdoniini
Genera Barberiella, Closterocoris, Cyphopelta, Dacerla, Heidemanniella, Mexicomiris, Paradacerla, Paraxenetus

Tribe Mirini

Tribe Resthenini
Genera Oncerometopus, Opistheurista, Platytylus, Prepops

Genera Acetropis, Actinocoris, Autumnimiris, Chaetofoveolocoris, Chaetomiris, Collaria, Dolichomiris, Leptopterna, Litomiris, Megaloceroea, Mimoceps, Pithanus, Porpomiris, Stenodema, Teratocoris, Trigonotylus

Subfamily Orthotylinae
Tribe Coridromiini(7)
Genera Coridromius

Genera Ceratocapsidea(8), Ceratocapsus, Pamillia, Pilophoropsidea(8), Pilophoropsis, Renodaeus(8), Schaffneria, Sericophanes

Tribe Halticini

Genera Acaciacoris, Aoplonema, Aoplonemella, Apachemiris, Argyrocoris, Ballella, Blepharidopterus, Brachynotocoris, Brooksetta, Cyrtorhinus, Daleapidea, Diaphnidia, Dichaetocoris, Ephedrodoma, Guerrerocoris(9), Fieberocapsus, Hadronema, Heterocordylus, Heterotoma, Ilnacora, Ilnacorella, Labopella, Labopidea, Lopidea, Lopidella, Mecomma, Melymacra, Noctuocoris, Origonema, Orthotylus, Parthenicus, Presidiomiris, Proboscidotylus, Pseudoloxops, Pseudopsallus, Pseudothenicus, Pseudoxenetus, Reuteria, Saileria, Scalponotatus, Slaterocoris, Squamocoris, Texocoris

Subfamily PHYLINAE
Genera Cyrtopeltocoris, Phoradendrepulus

Tribe Decomini
Genera Aurantiocoris

Tribe Nasocorini
Genera Adenostomocoris, Arctostaphylocoris, Atomoscelis, Atractotomus, Beckocoris, Campylomma, Chlamydatus, Chlamyopsallus, Hambletoniola, Knightomiroides, Larinocerus, Lattinophylus, Megalopsallus, Monosynamma, Nevadocoris, Phoenicocoris, Pinomiris, Pruneocoris, Rhinacloa, Spanagonicus, Tannerocoris

Tribe Exaeretini
Genera Gonoporomiris, Hyalopsallus, Tuponia

Subtribe Cremnorrhinina

Tribe Phylini
Subtribe Keltoniina

Subtribe Phylina
Genera Conostethus, Icodema, Izyaius, Lepidargyrus, Phylus, Psallus, Sthenarus

Subtribe Oncotylina
Genera Americodema, Arizonapsallus, Asciodema, Bisulcopsallus, Cariniocoris, Ceratopsallus, Compsidolon, Coniferocoris(10), Crassomiris, Europiella, Galbinocoris, Hamatophylus, Hoplomachus, Insulaphylus, Knightopiella, Knightopsallus, Lineatopsallus, Maculamiris, Megalocoleus, Occidentodema, Oligotylus, Oncotylus, Parapsallus, Phallospinophylus, Phyllopidea, Phymatopsallus, Piceophylus, Pinophylus, Plagiognathus, Plesiodema, Psallovius, Pygovepres, Quercophylus, Quernocoris, Ranzovius, Rhinocapsus, Rubellomiris, Rubeospineus, Salicopsallus, Sthenaropsidea, Stictopsallus, Vanduzeephylus, Vesperocoris

Tribe Semiini
Subtribe Semiina

Genera Alepidiella, Pilophorus, Sthenaridea

Subtribe Tuxedoina
Genera Tuxedo

*Proboscidotylus: first NA record in(11)
*Melymacra described in(12)
*Adenostomocoris, Arctostaphylocoris, Aurantiocoris, Guentherocoris, Pruneocoris, and Vanduzeephylus described in(13)
*Knightopiella described in(14)
*Calidroides and Chlamyopsallus described in(15)
* Izyaius described in(10)
*Galbinocoris, Hamatophylus, Insulaphylus, Maculamiris, Quernocoris, and Vesperocoris described in(16)
*Crassomiris, Phallospinophylus, Quercophylus, Pygovepres, Rubellomiris, and Rubeospineus described in(17)
*Arizonapsallus, Bisulcopsallus, Ceratopsallus, Cercocarpopsallus, Knightopsallus, Salicopsallus, and Stictopsallus described in(18)
*Origonema described in(19)
*Lattinophylus described in(20)
*Ambracius: first NA record in(21)
*Tytthus moved from the Leucophoropterini to Phylini sensu lato(22)
*Leutiola and Ticua described in(23)
*Phylinae is re-classified in(24)
Print References
Knight, H. H. 1968. Taxonomic review: Miridae of the Nevada Test Site and the western United States. Brigham Young University Science Bulletin 9(3): 1–282. (Biodiversity Heritage Library)
Knight, H. 1941. The Plant Bugs, or Miridae, of Illinois. Illinois Natural History Survey Bulletin 22, no. 1, 234 pp. (Illinois Natural History Survey)
Internet References
Miridae of Texas, compilation and crude sorting of the better TX mirid photos posted to BG (Quinn 2015)
The plant bugs of Denmark(25)
Works Cited
1.American Insects: A Handbook of the Insects of America North of Mexico
Ross H. Arnett. 2000. CRC Press.
2.Plant Bug Planetary Biodiversity Inventory
3.Biodiversity of the Heteroptera
Henry T.J. 2009. In: Foottit R.G., Adler P.H., eds. Insect biodiversity: Science and society. Oxford: Wiley-Blackwell: 223-263.
4.Schuh, R.T. 2002-2013. On-line Systematic Catalog of Plant Bugs (Insecta: Heteroptera: Miridae).
5.Review of the Eurychilopterella complex of genera, including the description of a new genus from Mexico
Gary M. Stonedahl, J. D. Lattin and V. Razafimahatrat. 1997. American Museum of Natural History.
6.Resurrection of the plant bug genus Pappus Distant, with Clarification of included species (Hemiptera: Heteroptera: Miridae)
Thomas J. Henry. 2006. Entomological Society of Washington.
7.Systematics, Biodiversity, Biogeography, and Host Associations of the Miridae (Insecta: Hemiptera: Heteroptera: Cimicomorpha)
G. Cassis and Randall T. Schuh. 2012. Annual Reviews.
8.Revision of the Ceratocapsine Renodaeus group: Marinonicoris, Pilophoropsis, Renodaeus, and Zanchisme, with...
Henry, Thomas J. 2015. Pensoft Publishers.
9.Revision and phylogenetic analysis of the North American genus Slaterocoris Wagner with new synonymy, ...
Michael D. Schwartz. 2011. American Museum of Natural History.
10.Review of Plesiodema Reuter and a description of a new genus to accommodate Psallus sericeus Heidemann ...
Michael D. Schwartz. 2006. KMK Scientific Press.
11.Proboscidotylus nigrosquamis (Maldonado) (Heteroptera:Miridae:Orthotylinae): New combination and first record for the United
Thomas J. Henry. 2003. Entomological Society of Washington.
12.Melymacra, a new plant bug genus (Heteroptera: Miridae: Orthotylinae) inhabiting manzanita, Arctostaphylos Adanson (Ericaceae)
Michael D. Schwartz. 2004. Pacific Coast Entomological Society.
13.New genera, new species, new synonyms, and new combinations in North America and Caribbean Phylinae (Heteroptera, Miridae)
Randall T. Schuh and Michael D. Schwartz. 2004. American Museum of Natural History.
14.Revision of Europiella Reuter in North America, with the Description of a New Genus (Heteroptera: Miridae: Phylinae)
Randall T. Schuh. 2004. American Museum of Natural History.
15.Redescription of Strophopoda aprica Van Duzee and the Description of Two New Genera and Five New Species from ...
Michael D. Schwartz. 2005. American Museum of Natural History.
16.New genera, new species, and new combinations in western Nearctic Phylini (Heteroptera, Miridae, Phylinae)
Christiane Weirauch. 2006. American Museum of Natural History.
17.New Genera and Species of Oak-Associated Phylini (Heteroptera: Miridae: Phylinae) from Western North America
Christiane Weirauch. 2006. American Museum of Natural History.
18.Revision, phylogenetic, biogeographic, and host analysis of the endemic western North American Phymatopsallus group, ...
Randall T. Schuh. 2006. American Museum of Natural History.
19.Revision and phylogenetic analysis of the Hadronema Group (Miridae, Orthotylinae, Orthotylini), ...
Dimitri Forero. 2008. American Museum of Natural History.
20.Beckocoris Knight, Nevadocoris Knight, and Lattincoris artemisii, new genus and new species, Asteraceae feeding Phylinae
Randall T. Schuh. 2008. Entomological Society of Washington.
21.Revision of the genus Ambracius Stål, 1860 (Heteroptera: Miridae: Deraeocorinae: Clivinematini), with descriptions of ...
Paulo Sérgio F Ferreia & Thomas J. Henry. 2010. Magnolia Press.
22.Revision of Leucophoropterini: ...
Katrina L. Menard & Randall T. Schuh. 2011. American Museum of Natural History.
23.Revision of the Nearctic Genus Coquillettia Uhler With a Transfer to the Tribe Phylini, the Description of 14 New Species, ...
Denise Wyniger. 2011. New York Entomological Society.
24.A revised classification of the Phylinae (Insecta, Heteroptera, Miridae) : arguments for the placement of genera
Randall T. Schuh and Katrina L Menard. 2013. American Museum of Natural History.
25.Skipper L. (2003-2012) Danmarks Blomstertæger