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Photo#971703
Bee fly on chorizanthe - Lepidanthrax

Bee fly on chorizanthe - Lepidanthrax
Fort Ord, Monterey County, California, USA
June 11, 2014
I photographed at least two individuals of Lepidanthrax visiting these flowers of Chorizanthe diffusa on this summer morning. Another individual appears in the post below:



...and Geminaria canalis was also seen on these flowers.

Using the Bombyliidae key in the MND(1) I originally erred in keying this to genus Paravilla (not noticing the long proboscis here, and getting derailed at couplet 54). Thankfully, Joel Kits noted my error and saw this was Lepidanthrax. The corrected couplet sequence from the MND(1) for keying to genus here appears below:

 1) Hind margin of eye sharply and deeply indented....38
38) R2+3, arising from R4+5, near crossvein r-m, and at a right angle.....45
45) First flagellomere with an apical stylus (and without an apical tuft of hair)......47
47) Stylus not distinctly articulated with first flagellomere......49
49) Veins CuA2 and A ending separately in wing margin.......50
50) R2+3 bent apically but not strongly contorted, not connected to R4 by a crossvein.....53
53) Veins CuA2 and A converging toward wing margin...........54
54) Proboscis strongly projecting beyond subcranial margin.....55
55) R2+3, and R4 not connected by a crossvein (Fig. 44).....56
56) At least basal half of abdomen with broad scales; male usually with apex of abdomen silvery. Fore tibia with rows of short bristles. Crossvein dm-cu not much longer than middle portion of CuA, (Fig. 44)........Lepidanthrax

The characters at couplets 45 and 47 are not clearly visible in my photos here, but the alternatives they lead to can be eliminated further along in the key by noting various characters conflicts that arise.

Images of this individual: tag all
Bee fly on chorizanthe - Lepidanthrax Bee fly on chorizanthe - Lepidanthrax Bee fly on chorizanthe - Lepidanthrax

Moved
These are actually Lepidanthrax. The long proboscis isn't obvious in these photos (the base is visible here, extending into the flower), so the key fails at that point.

Moved from Paravilla.

 
Thanks for the correction, Joel!
I very much appreciate your reviewing my images and notes and making corrections of errors...and am especially grateful you were able to indicate the point where my analysis went astray at couplet 54 (e.g. the projecting proboscis here). Would have liked to respond earlier, but wanted to study the MND key and Hall's 1976 Lepidanthrax paper first, and have been too busy with work to do so up to now.

I do see the conical base of the proboscis now, with a short thin extension...a bit longer than the conical base...with its tip appearing to touch a flat, white, perianth lobe. And it became clear to me that this goes to Lepidanthrax, once you pointed me in the right direction at couplet 54 in the MND.

I tried to work through Hall(1)(1976), but couldn't make clear progress...I can't tell whether this is a male or a female, which is the first step Hall's keys. Going each way, I tentatively get to either the angulus group or the campestris group. Going through the key and descriptions in Hall for members of the angulus group, nothing seemed to fit well. On the other hand, many parts of the description of L. campestris seem to fit, and the Monterey locality here is within range. But I'm not at all confident of that species ID.

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