Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes

Upcoming Events

Information about the 2019 BugGuide Gathering in Louisiana, July 25-27

Discussion, insects and people from the 2018 gathering in Virginia, July 27-29

Photos of insects and people from the 2015 gathering in Wisconsin, July 10-12

Photos of insects and people from the 2014 gathering in Virginia, June 4-7.

Photos of insects and people from the 2013 gathering in Arizona, July 25-28

Photos of insects and people from the 2012 gathering in Alabama

Photos of insects and people from the 2011 gathering in Iowa

Revision of Phanaeus MacLeay, A New World genus of scarabaeine dung beetles
By Edmonds, W.D. 1994
Natural History Museum of Los Angeles County, Contributions to Science 443, 1994
Cite: 9734
Beautifully illustrated revision of this very popular group

Full PDF (in B&W) - Discussion of VINDEX group starts on page 83.

Edmonds, W.D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeine dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Natural History Museum of Los Angeles County, Contributions in Science 443: 1-105, 360 figs.

ABSTRACT. This study is a revision of the genus Phanaeus as defined by Edmonds (1972). The introduction includes a history of the genus, a key separating it from related genera, a discussion of taxonomic characters and primary type material, a summary of the biology and geography of the genus, and conclusions about its evolution. The systematic section treats 48 species and subspecies in two subgenera, one of which, Notiophanaeus, is new. Notiophanaeus comprises five species groups embracing 15 mostly South American species. Phanaeus, sen. str., includes 31 species and subspecies arranged in eight species groups. Keys separate subgenera, species groups, species, and subspecies. [snip]

A list of all known available names and a previously established synonymy are included in the introduction. Geographical distribution data for all species and subspecies are contained in the appendix.

Comment viewing options
Select your preferred way to display the comments and click 'Save settings' to activate your changes.